|
There is
a wide variety of wetland habitats for the naturalist
willing to get wet in the pursuit of new species and
unique habitats. Several grassland communities are the
most diverse sites in the region, with strong floristic
ties to Northwestern hot spots in the Olympic Mountains
and Columbia Gorge.
Coniferous
forest is the dominant habitat of the coast, and within
this category, industrial timberland is the most common
landuse category. Commercial timber aside, the area
still has a few stands of old growth trees.
Summer drought is the reason that conifers dominate
many western forests of North America. Despite wet winters,
dry growing seasons along the Northwest Coast severely
limit the spread of broad-leaved deciduous trees. Here
hardwoods grow only in perennially damp soils. Grasslands
are restricted to even drier sites. Only conifers flourish
in wet winters and dry summers.
Dynamics
Plant communities develop through succession, involving
change in species composition over time. Coast range
forests north of the Nehalem River in Oregon developed
without regular fire, but with regular severe windstorms.
Consequently these forests exhibit highly diverse age
structure with complex, multilayered
understories and a few dominant tree species. Wind throw
was uncommon over large areas prior to the advent of
industrial forestry. Shade tolerance characterizes the
dominant tree species, allowing seedlings to establish
in the shade of mature trees and wait for wind to top
or topple them, creating light gaps that release the
young trees and allow them to grow.
Wildfires, when they occur in these forests, are catastrophic,
burning thousands of acres at a time. Restoration of
conifer overstories may take fifty to one hundred years
or more. Dominant conifers are thin-barked and not fire
tolerant. Seeds are not fire-tolerant. Fires burn the
forest duff layer, a thick layer of slowly decomposing
wood, needles and bark on the forest floor that holds
nutrients and water, and supports soil fungi. Elimination
of duff by wildfire may seriously slow forest restoration,
as soil fungi are critical partners to conifer seedlings.
Coastal
Temperate Rainforest
The coastal temperate rainforest of North America is
the most extensive example of this forest type in the
world. Found only on the West Coast, coastal temperate
rainforest occurs from Northern California near San
Francisco Bay to South-central Alaska, growing in a
narrow fringe, with a small parallel fringe on the west
side of the Cascade Range from central Oregon to southern
British Columbia. Temperate rainforests are distinguished
by rarity of fire, evergreeness, and complex physical
structure (abundance of epiphytes, multiple canopy layers
under the main canopy, and a dense understory).
Dense coniferous forests are highly productive. Throughout
the Columbia Coast, cleared patches readily return to
dense forest within fifty years. Shrubby understories
are legendary. Accustomed to Eastern hardwood forests,
an archeologoist was hiking on Long Island in the 1970s,
looking for shell middens. In his frustration at the
dense, cloaking vegetation, he uttered what could be
a classic statement about coastal vegetation, "You
could put the White House here, walk away from it for
twenty years, and we'd never find it again."
Ancient Forests
"Productive, low-elevation, old-growth temperate
rainforests are among the most massive ecosystems on
earth. Tremendous accumulations of biomass result from
the combination of: long periods of time between stand-destroying
disturbances; the great longevity of many northwestern
trees; their inherent ability to grow large; and relatively
slow decomposition rates. Frequently more than half
of the total mass in these forests is in the form of
dead trees, either snags or logs. . . The great abundance
of dead, woody material in such forests has led to the
development of complex communities of organisms that
depend on decomposing material . . . structural attributes
characteristic of older forests are a wide range of
tree sizes and ages, and a patchy, open canopy punctuated
by gaps beneath which the forest understory is especially
well developed." Quote from Plants of the Pacific
Northwest Coast, used with permission by the publisher.
Individual
tree ages of 600 to >1,000 years are not uncommon
in ancient forests. Growth rates vary widely by site;
size cannot be used to establish tree age. Sitka spruce
trees along the ocean on wet clay soils and more than
five feet dbh may be less than 200 years old, while
spruces of the same diameter growing 8-10 miles inland
may be more than 600 years old.
|